5.0 liter tracing its roots to a Ford Racing engine.
From what I understand though their new motors are completely their own design. I'm waiting for their next motor which is supposed to eliminate the cam shafts and use a form of valve train technology seen in F1 using pneumatic actuators.
As it sits, the Agera R is and incredible machine. Here's a little video of the factory where these demons are incarnated into automotive greatness: